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Malaria

Infectious Diseases · Part 5 – Infectious Diseases: Parasitic · Part 5 – Infectious Diseases: Parasitic · Chapter 231


Key Clinical Points

  1. Malaria is a protozoan disease transmitted by infected female Anopheles mosquitoes and is the most important parasitic disease of humans.
  2. Six Plasmodium species cause human infections: P. falciparum, P. vivax, P. ovale (curtisi/wallikeri), P. malariae, and P. knowlesi.
  3. P. falciparum predominates in Africa, New Guinea, and Hispaniola; P. vivax is more common in Central/South America and Asia.
  4. Severe falciparum malaria involves sequestration of infected RBCs in vital organs via PfEMP1-mediated cytoadherence.
  5. Cerebral malaria presents as diffuse symmetric encephalopathy with rare focal signs.
  6. Hypoglycemia is a common complication in severe malaria, especially in children and pregnant women.
  7. Malaria in early pregnancy causes fetal loss; falciparum malaria in primigravid women correlates with low birth weight.
  8. Most falciparum malaria deaths (~600,000/year) occur in young African children.
  9. Clinical diagnosis is unreliable; classic paroxysms are rare and suggest relapse with P. vivax/ovale.
  10. Distinct parasite morphologies (e.g., banana-shaped gametocytes for P. falciparum, Schüffner's dots for P. vivax) are critical for identification.

1. DEFINITION & OVERVIEW

Malaria is a protozoan disease transmitted by the bite of infected female Anopheles mosquitoes and is the most important of the parasitic diseases of humans.

Historical Context: • Elimination from temperate zones (US, Canada, Europe, Russia) >50 years ago • Prevalence rose in tropics 1970–2000 • Mortality reduction 2000–2015 due to control programs • Resurgence since 2015 with rising case numbers in sub-Saharan Africa • Elimination efforts hindered by drug/insecticide resistance


2. EPIDEMIOLOGY

Global Burden: • ~249 million cases and 608,000 deaths globally in 2022 • ~1,660 deaths daily

Species Distribution: • P. falciparum: Africa, New Guinea, Hispaniola • P. vivax: Central/South America, Asia • P. ovale: Africa (curtisi/wallikeri) • P. malariae: Sub-Saharan Africa and globally • P. knowlesi: Borneo, Southeast Asia

Endemicity Classification: • Hypoendemic: <1% parasitemia in children 2–9 years • Mesendemic: 1–75% • Holoendemic: >75%

Vector Biology: • Transmission proportional to vector density, human-bite rate², and mosquito survival • Sporogony requires >7-day mosquito survival (8–30 days depending on temperature)


3. ETIOLOGY & PATHOPHYSIOLOGY

Parasite Characteristics (Table 1): • P. falciparum: 5.5-day intrahepatic phase; 30,000 merozoites released; 48h erythrocytic cycle; infects younger cells; no relapses; morphology includes ring forms and banana-shaped gametocytes. • P. vivax: 8-day intrahepatic phase; 10,000 merozoites released; 48h erythrocytic cycle; infects reticulocytes/cells up to 2 weeks old; relapses possible; morphology includes large rings and Schüffner's dots. • P. ovale: 9-day intrahepatic phase; 15,000 merozoites released; 50h erythrocytic cycle; infects reticulocytes; relapses possible; morphology includes enlarged erythrocytes with tufted ends. • P. malariae: 15-day intrahepatic phase; 15,000 merozoites released; 72h erythrocytic cycle; infects older cells; no relapses; morphology includes band or rectangular forms. • P. knowlesi: 5.5-day intrahepatic phase; 20,000 merozoites released; 24h erythrocytic cycle; infects younger cells; no relapses; resembles P. falciparum/P. malariae.

Pathogenesis of Severe Falciparum Malaria: • PfEMP1-mediated cytoadherence to endothelial receptors (ICAM-1, EPCR) • Sequestration in brain, lungs, kidneys → microvascular obstruction • Rosetting/agglutination of infected RBCs → exacerbates sequestration • Asymptomatic parasitemia common in endemic areas (~5000/mL) • Immunity: Developed through strain-specific IgM/IgG responses

Immune Targets (Figure 1)

Liver Stage: CD4+ and CD8+ T cells kill intrahepatic parasites • Blood Entry: Antibodies to merozoites block invasion of RBCs • Toxins: Antibodies to malaria 'toxins' • Coadherence/Clearance: Antibodies to parasite antigens on infected RBCs block cytoadherence to endothelium and augment splenic clearance • Mosquito Stage: Antibodies block fertilization, development, and invasion


4. CLINICAL FEATURES

General Presentation: • Early phase: Malaise, headache, myalgia, abdominal pain • Severe cases: Hyperpyrexia (>40°C), tachycardia, delirium

Severe Falciparum Malaria (Table 2):Cerebral Malaria: Unarousable coma; failure to respond to stimuli; duration >30 min after seizure; GCS ≤8 (or Blantyre <3 in children) • Acidemia/Acidosis: Arterial pH ≤7.3, bicarbonate ≤15 mmol/L; venous lactate ≥5 mmol/L • Severe Anemia: Hemoglobin <50 g/L (<5 g/dL) with parasite density >100,000/μL • Renal Failure: Creatinine >265 μmol/L (>3 mg/dL) or BUN >20 mmol/L • Pulmonary Edema/ARDS: Noncardiogenic; oxygen saturation <92% on room air; RR >30/min • Hypoxemia: Plasma oxygen content <60 mL/dL • Hypotension/Shock: Systolic BP <70 mmHg (children 1–5y) or <80 mmHg (adults); capillary refill ≥2 s • Bleeding/DIC: Significant hemorrhage in gums, nose, GI tract; evidence of DIC • Convulsions: >2 generalized seizures in 24 h • Hyperparasitemia: >5% in nonimmune patients (≥10% in any patient) • Jaundice: Bilirubin >34 μmol/L (>3 mg/dL) with parasite density >100,000/μL

Other Clinical Findings: • Hemoglobinuria: Macroscopic black, brown, or red urine • Extreme weakness: Prostration; inability to sit unaided • Ocular signs (Figure 3): Perimacular whitening and pale-centered retinal hemorrhages


5. DIFFERENTIAL DIAGNOSIS

Viral illness: Fever without rash, no meningismus • Dengue: Less severe myalgia, no hemorrhagic manifestations • Typhoid: No abdominal tenderness, no rose spots • Leptospirosis: No jaundice in uncomplicated cases • Bacterial meningitis: No neck stiffness, no purulent CSF


6. INVESTIGATIONS & DIAGNOSIS

  1. Clinical Suspicion: • Fever in endemic area with travel history • History of recent antimalarial drug use • Presence of splenomegaly or jaundice
  2. Microscopy (Table 4): • Thick film: Used for parasite quantification; sensitive (0.001% parasitemia); identifies species • Thin film: Used for morphology and specific identification of stages (e.g., ring forms, trophozoites)
  3. Rapid Diagnostic Tests (RDTs): • HRP-2 (P. falciparum), pLDH (all species), aldolase • Advantage: Rapid; sensitivity similar to thick films (~0.001% parasitemia)
  4. Molecular Methods: • PCR for species confirmation and mixed infections
  5. Specialized Techniques: • Microtube concentration with acridine orange (high sensitivity, but no speciation)

7. MANAGEMENT & TREATMENT

  1. Uncomplicated Malaria (Table 5): • Chloroquine: For known chloroquine-sensitive strains; 10 mg/kg stat followed by 5 mg/kg at 12, 24, and 36 h OR 10 mg/kg at 24 h and 5 mg/kg at 48 h • Amodiaquine: 10–12 mg/kg qd for 3 days
  2. Radical Treatment (Prevention of Relapse): • Primaquine: For P. vivax or P. ovale; 0.5 mg/kg qd for 14 days (Southeast Asia/Oceania) or 0.25 mg/kg elsewhere • Tafenoquine: Single dose 300 mg (only if G6PD levels are normal)
  3. Severe Falciparum Malaria: • Artesunate: 4 mg/kg qd for 3 days • Combination options:
  4. Artesunate + Sulfadoxine (25 mg/kg) / Pyrimethamine (1.25 mg/kg)
  5. Artesunate + Amodiaquine (10 mg/kg qd for 3 days)
  6. Artemether-lumefantrined (1.5/9 mg/kg bid for 3 days with food)
  7. DHA-piperaquined (target dose: 4/24 mg/kg qd for 3 days in children <25 kg; 4/18 mg/kg qd for 3 days in persons ≥25 kg)
  8. Artesunate-pyronaridine (4/12 mg/kg qd for 3 days)
  9. Support: Exchange transfusion for severe anemia (Hb <50 g/L); Dialysis for AKI
  10. Prophylaxis (Table 8):
  11. Atovaquone-proguanil (Malarone): 1 adult tablet daily; 300 mg base once weekly (specific cases). Must be taken with food or a milky drink.
  12. Doxycycline: 100 mg qd (adult) or 2 mg/kg qd (pediatric ≥8 years)
  13. Mefloquine: 228 mg base once weekly; 30 mg base qd for P. vivax
  14. Primaquine: 30 mg base qd for 14 days post-exposure

8. PROGNOSIS & COMPLICATIONS

Acute Complications: • Cerebral malaria: Mortality 15–20%; long-term neurological sequelae (epilepsy, cognitive deficits) • ARDS: Seen in 5–10% of cases • AKI: Creatinine >3 mg/dL • Severe anemia: Hb <5 g/dL

Pregnancy Complications: • Maternal mortality risk increased 3–5x • Fetal loss in first trimester; low birth weight in second/third • Placental sequestration with VAR2CSA binding

Pediatric Complications: • Higher parasite density threshold for symptoms (~100,000/μL) • Increased risk of cerebral malaria and anemia • Febrile convulsions (5–10% of cases)

Risk Factors (Table 3): • Anemia: Very frequent in children (+++) • Hypoglycemia: Very frequent in pregnant women (+++) and children (+++) • Renal failure: Frequent in nonpregnant adults (+++) and pregnant women (+++) • Pulmonary edema: Frequent in nonpregnant adults (+++) and pregnant women (+++)


9. SPECIAL CONSIDERATIONS

Pregnancy: • Increased risk of maternal mortality (3–5x higher) • Fetal loss in first trimester, low birth weight in second/third • Placental sequestration with VAR2CSA binding

Children: • Higher parasite density thresholds for symptoms (~100,000/μL) • Increased risk of cerebral malaria and anemia • Febrile convulsions common (5–10% of cases)


10. KEY PEARLS & CLINICAL TRAPS

Diagnostic Pitfalls: • Reliance on clinical diagnosis (sensitivity <30%) • Atypical presentations in immunocompromised patients • Mixed infections with multiple species

Prognostic Indicators (Table 2): • Platelet count <50,000/μL predicts poor outcome • Creatinine >265 μmol/L (>3 mg/dL) indicates severe AKI • Bilirubin >34 μmol/L (>3 mg/dL) with parasite density >100,000/μL • Hyperparasitemia (>5% in nonimmune patients; ≥10% in any patient)

Drug Specifics (Table 6): • Quinine: Causes cinchonism; acts on trophozoite blood stage • Primaquine: Risk of severe hemolytic anemia in G6PD deficiency • Artemether/Artesunate: Rapidly eliminated; no action on liver stages


Reference Tables

TABLE 231-1 Characteristics of Plasmodium Species Infecting Humans CHARACTERISTIC Duration of intrahepatic phase (days)…

Harrison's 22e, p.1760

CHARACTERISTIC FINDING FOR INDICATED SPECIES
P. FALCIPARUM P. VIVAX P. OVALEa P. MALARIAE P. KNOWLESI
Duration of intrahepatic phase (days) 5.5 8 9 15 5.5
30,000 10,000 15,000 15,000
Approximate duration of erythrocytic
cycle (h)
48 48 50 72 24
Younger cells (but can
invade cells of all ages)
Reticulocytes and cells
up to 2 weeks old
Reticulocytes Older cells
Morphology Usually only ring
forms; banana-shaped
gametocytes
Irregularly shaped large
rings and trophozoites;
enlarged erythrocytes;
Schüffner’s dots
Infected erythrocytes,
enlarged and oval
with tufted ends;
Schüffner’sb dots
Band or
rectangular forms
of trophozoites
common
Resembles P. falciparum (early
trophozoites) or P. malariae
(later trophozoites, including
band forms)
Black Yellow-brown Dark brown Brown-black
Ability to cause relapses No Yes Yes No No

TABLE 231-3 Features Indicating a Poor Prognosis in Severe Falciparum Malaria Clinical Marked agitation…

Harrison's 22e, p.1764

SIGNS MANIFESTATIONS
Major
Unarousable coma/
cerebral malaria
Failure to localize or respond appropriately to noxious
stimuli; coma persisting for >30 min after generalized
convulsion; a Glasgow Coma Score <11, or in young
children a Blantyre Coma Score of <3

TABLE 231-3 Features Indicating a Poor Prognosis in Severe Falciparum Malaria

Clinical
Marked agitation
Hyperventilation (respiratory distress)
Low core temperature (<36.5°C; <97.7°F)
Bleeding
Deep coma
Repeated convulsions
Anuria
Shock
Laboratory
Biochemistry
Hypoglycemia (<2.2 mmol/L)
Hyperlactatemia (≥5 mmol/L)
Acidemia (arterial pH <7.25, base deficit >8 meq/L, or serum HCO <15 mmol/L)
3
Elevated serum creatinine (>265 μmol/L)
Elevated total bilirubin (>50 μmol/L)
Elevated liver enzymes (AST/ALT 3 times upper limit of normal)
Elevated muscle enzymes (CPK ↑, myoglobin ↑)
Elevated urate (>600 μmol/L)
Hematology
Leukocytosis (>12,000/μL)
Severe anemia (PCV <10%)
Coagulopathy
Low platelet count (<50,000/μL)
Prolonged prothrombin time (>3 s)
Prolonged partial thromboplastin time
Decreased fibrinogen (<200 mg/dL)
Parasitology
Hyperparasitemia
Increased mortality at >100,000/μL
High mortality at >500,000/μL
>20% of parasites identified as pigment-containing trophozoites and schizonts
>5% of neutrophils contain visible malaria pigment
Severe
normochromic,
normocytic anemia
Hematocrit of <15% or hemoglobin level of <50 g/L (<5 g/dL)
with parasite density of >10,000/μLa
Pulmonary edema/
adult respiratory
distress syndrome
Noncardiogenic pulmonary edema, often aggravated
by overhydration; radiologically confirmed or oxygen
saturation <92% on room air with a respiratory rate >30/
min, often with chest wall indrawing and crepitations on
auscultation
Hypotension/shock Systolic blood pressure of <70 mmHg in children 1–5 years or
<80 mmHg in adults; with evidence of impaired perfusion
or capillary refill >2 s
Convulsions More than two generalized seizures in 24 h; signs of
continued seizure activity, sometimes subtle (e.g., tonic-
clonic eye movements without limb or face movement)
Other
Extreme weakness Prostration; inability to sit unaidedd
Jaundice Serum bilirubin level of >50 mmol/L (>3 mg/dL) if combined
with a parasite density of >100,000/μL or other evidence
of vital-organ dysfunction

TABLE 231-4 Relative Incidence of Severe Complications of Falciparum Malaria COMPLICATION Anemia Convulsions…

Harrison's 22e, p.1765

COMPLICATION NONPREGNANT
ADULTS
PREGNANT
WOMEN
CHILDREN
Anemia + ++ +++
+ +
Hypoglycemia + +++ +++
+++ +++
Renal failure +++ +++ +
++ +++

TABLE 231-5 Standard Methods for the Diagnosis of Malaria a METHOD Thick blood film b

Harrison's 22e, p.1769

METHOD PROCEDURE ADVANTAGES DISADVANTAGES
Thick blood filmb Blood should be uneven in thickness but thin enough that the hands
of a watch can be read through part of the spot. Stain dried, unfixed
blood spot with Giemsa, Field’s, or another Romanowsky stain.
Count number of asexual parasites per 200 WBCs (or per 500 WBCs
at low densities). Count and report gametocytes separately.c
Sensitive (0.001% parasitemia);
species specific; inexpensive
Requires experience (artifacts may be
misinterpreted as low-level parasitemia);
underestimates true count
Stain fixed smear with Giemsa, Field’s, or another Romanowsky
stain. Count number of RBCs containing asexual parasites per 1000
RBCs. In severe malaria, assess stage of parasite development and
count neutrophils containing malaria pigment.e Count and report
gametocytes separately.c
Rapid; species specific;
inexpensive; in severe
malaria, provides prognostic
informatione
PfHRP2 dipstick
or card test
A drop of blood is placed on the stick or card, which is then
immersed in washing solutions. Monoclonal antibody capture of
parasitic antigens reads out as a colored band.
Robust and relatively
inexpensive; rapid; sensitivity
similar to or slightly lower than
that of thick films (~0.001%
parasitemia)
Detects only Plasmodium falciparum;
remains positive for weeks after high-density
infectionsf; does not quantitate P. falciparum
parasitemia; evasion of detection by certain
strains due to polymorphisms/deletions in
HRP2/3 genes
A drop of blood is placed on the stick or card, which is then
immersed in washing solutions. Monoclonal antibody capture of
parasitic antigens reads out as two colored bands. One band is
genus specific (all malarias) or P. vivax specific, and the other band
is specific for P. falciparum.
Rapid; sensitivity similar to or
slightly lower than that of thick
films for P. falciparum (~0.001%
parasitemia)
Microtube
concentration
methods with
acridine orange
staining
Blood is collected in a specialized tube containing acridine orange,
anticoagulant, and a float. After centrifugation, which concentrates
the parasitized cells around the float, fluorescence microscopy is
performed.
Sensitivity similar or superior
to that of thick films (~0.001%
parasitemia); ideal for
processing large numbers of
samples rapidly
Does not speciate or quantitate; requires
fluorescence microscopy

TABLE 231-6 Regimens for the Treatment of Malaria a

Harrison's 22e, p.1770

TYPE OF DISEASE OR TREATMENT REGIMEN(S)
Uncomplicated Malaria
Known chloroquine-sensitive strains of
Plasmodium vivax, P. malariae, P. ovale,
P. falciparumb
Chloroquine (10 mg of base/kg stat followed by 5 mg/kg at 12, 24, and 36 h or by 10 mg/kg at 24 h and 5 mg/kg at 48 h)
or
Amodiaquine (10–12 mg of base/kg qd for 3 days)
Radical treatment for P. vivax or
P. ovale infection (prevention of
relapse)
In addition to chloroquine or amodiaquine or ACT, primaquine (0.5 mg of base/kg qd in Southeast Asia and Oceania [total
dose 7 mg/kg] and 0.25 mg/kg elsewhere [total dose 3.5 mg/kg]) should be given for 14 days to prevent relapse.c In mild
G6PD deficiency, 0.75 mg of base/kg should be given once weekly for 8 weeks. Primaquine should not be given in severe
G6PD deficiency. Single dose tafenoquine (adult dose 300 mg) is being introduced for radical cure in some areas alongside
quantitative G6PD testing. Can be given only If G6PD levels are normal.
P. falciparum malaria Artesunated,e (4 mg/kg qd for 3 days) plus sulfadoxine (25 mg/kg)/pyrimethamine (1.25 mg/kg) as a single dose
or
Artesunated (4 mg/kg qd for 3 days) plus amodiaquine (10 mg of base/kg qd for 3 days)d,e
or
Artemether-lumefantrined (1.5/9 mg/kg bid for 3 days with food)
or
Artesunated (4 mg/kg qd for 3 days) plus mefloquine (24–25 mg of base/kg—either 8 mg/kg qd for 3 days or 15 mg/kg on day 2
and then 10 mg/kg on day 3)f
or
DHA-piperaquined (target dose: 4/24 mg/kg qd for 3 days in children weighing <25 kg and 4/18 mg/kg qd for 3 days in persons
weighing ≥25 kg)
or
Artesunate-pyronaridined (4/12 mg/kg qd for 3 days)
Second-line treatment/treatment of
imported malaria (if other ACT not
available)
Artesunatee (2 mg/kg qd for 7 days) or quinine (10 mg of salt/kg tid for 7 days) plus 1 of the following 3:
Tetracyclinef (4 mg/kg qid for 7 days)
Doxycyclinef (3 mg/kg qd for 7 days)
Clindamycin (10 mg/kg bid for 7 days)
or
Atovaquone-proguanil (20/8 mg/kg qd for 3 days with food)
Severe Falciparum Malariag,h,i

TABLE 231-7 Properties of Antimalarial Drugs DRUG(S) a Quinine

Harrison's 22e, p.1771

DRUG(S)a PHARMACOKINETIC PROPERTIES ANTIMALARIAL ACTIVITY MINOR TOXICITY MAJOR TOXICITY
Quinine Good oral and IM absorption
(quinine); Cl and V reduced, but
d
plasma protein binding (principally
to α1 acid glycoprotein) increased
(90%) in malaria; quinine t : 16 h in
1/2
malaria, 11 h in healthy persons
Acts mainly on trophozoite blood stage;
kills gametocytes of P. vivax, P. ovale,
and P. malariae (but not P. falciparum);
no action on liver stages
Common: cinchonism (tinnitus,
high-tone hearing loss, nausea,
vomiting, dysphoria, postural
hypotension); ECG QT interval
prolongation (usually by
<10%). Rare: diarrhea, visual
disturbance, rashes. Note: very
bitter taste
Common: hypoglycemia.
Rare: hypotension, blindness,
deafness, cardiac arrhythmias,
thrombocytopenia, hemolysis,
hemolytic-uremic syndrome,
vasculitis, cholestatic hepatitis,
neuromuscular paralysis.
Good oral absorption, very rapid
IM and SC absorption; complex
pharmacokinetics; enormous Cl and
V (unaffected by malaria); blood
d
concentration profile determined by
distribution processes in malaria;
t : 1–2 months. Active desethyl
1/2
metabolite about 25% of parent drug
concentrations
As for quinine, but acts slightly earlier
in asexual cycle
Common: nausea, dysphoria,
pruritus in dark-skinned
patients, postural hypotension,
ECG QT prolongation. Rare:
accommodation difficulties,
keratopathy, hypoglycemia,
rash. Note: bitter taste but
usually well tolerated
Piperaquine Adequate oral absorption, may
be enhanced by fats; similar
pharmacokinetics to chloroquine; t :
1/2
21–28 days
As for chloroquine; resistance has
emerged in Southeast Asia
Occasional epigastric pain,
diarrhea, ECG QT prolongation
None identified
Good oral absorption; largely
converted to active metabolite
desethylamodiaquine; t : 4–5 days
1/2
As for chloroquine, but more active
against chloroquine-resistant
P. falciparum
Nausea (tastes better than
chloroquine), dysphoria,
headache, bradycardia, ECG QT
prolongation
Primaquine Complete oral absorption; active
metabolite produced mainly via
CYP2D6; t : 5–7 h
1/2
Radical cure; eradicates hepatic
forms of P. vivax and P. ovale; kills
P. falciparum gametocytes; kills
developing liver stages of all species
Nausea, vomiting, diarrhea,
abdominal pain, hemolysis,
methemoglobinemia
Serious hemolytic anemia
in severe G6PD deficiency;
hemoglobinuria
Adequate oral absorption; no
parenteral preparation; t :
1/2
14–20 days (shorter in malaria)
As for quinine; resistance has emerged
in Southeast Asia
Nausea, giddiness, dysphoria,
fuzzy thinking, sleeplessness,
nightmares, sense of
dissociation
Lumefantrine Highly variable absorption related to
fat intake; t : 3–4 days
1/2
As for quinine None identified None identified
Good oral absorption; good
absorption of IM artesunate but
slow and variable absorption of
IM artemether; artesunate and
artemether biotransformed to active
metabolite dihydroartemisinin; all
drugs eliminated very rapidly; t : <1 h
1/2
Broader stage specificity and more
rapid than other drugs; no action on
liver stages; kills all but fully mature
gametocytes of P. falciparum
Reduction in reticulocyte count
(but not anemia); neutropenia
at high doses; in some cases,
delayed anemia after treatment
of severe malaria with
hyperparasitemia
Pyrimethamine Good oral absorption, variable IM
absorption; t : 4 days
1/2
For blood stages, acts mainly on mature
forms; causal prophylactic
Well tolerated Megaloblastic anemia,
pancytopenia, pulmonary
infiltration
Good oral absorption; biotransformed
to active metabolite cycloguanil;
t : 16 h; biotransformation reduced
1/2
by oral contraceptive use and in
pregnancy
Causal prophylactic; not used alone for
treatment
Well tolerated; mouth ulcers
and rare alopecia
Atovaquoneb Highly variable absorption related to
fat intake; t : 30–70 h
1/2
Acts mainly on trophozoite blood stage None identified None identified
Excellent absorption; t : 8 h for
1/2
tetracycline, 18 h for doxycycline
Weak antimalarial activity; should not
be used alone for treatment
Gastrointestinal intolerance,
deposition in growing bones
and teeth (tetracycline),
photosensitivity, moniliasis,
benign intracranial
hypertension
Pyronaridine Rapid variable absorption, large V;
d
t : 12–14 days
1/2
Acts mainly on trophozoite blood stage;
kills gametocytes of P. vivax, P. ovale,
and P. malariae (but not P. falciparum);
no action on liver stages
Gastrointestinal intolerance,
anemia, transient elevation
of aminotransferases,
hypoglycemia, headache
None identified
t : 3 h
1/2
Broad stage specificity; no action on
liver stages; kills all but fully mature
gametocytes of P. falciparum
Gastrointestinal intolerance,
transient elevation of
aminotransferases

TABLE 231-8 Drugs Used in the Prophylaxis of Malaria a DRUG Atovaquone- proguanil (Malarone)

Harrison's 22e, p.1774

DRUG USAGE ADULT DOSE PEDIATRIC DOSE COMMENTS
Atovaquone-
proguanil (Malarone)
Prophylaxis in areas
with chloroquine- or
mefloquine-resistant
Plasmodium falciparum
1 adult tablet POb 5–8 kg: ½ pediatric tabletc daily
≥8–10 kg: ¾ pediatric tablet
daily
≥10–20 kg: 1 pediatric tablet
daily
≥20–30 kg: 2 pediatric tablets
daily
≥30–40 kg: 3 pediatric tablets
daily
≥40 kg: 1 adult tablet daily
Begin 1–2 days before travel to malarious
areas. Take daily at the same time each day
while in the malarious areas and for 7 days
after leaving such areas. Atovaquone-proguanil
is contraindicated in persons with severe
renal impairment (creatinine clearance rate,
<30 mL/min). In the absence of data, it is not
recommended for children weighing <5 kg,
pregnant women, or women breast-feeding
infants weighing <5 kg. Atovaquone-proguanil
should be taken with food or a milky drink.
Prophylaxis only in the
very few areas with
chloroquine-sensitive
P. falciparumc or areas
with P. vivax only
300 mg of base (500 mg of salt)
PO once weekly
5 mg of base/kg (8.3 mg of salt/
kg) PO once weekly, up to
maximum adult dose of 300 mg
of base
Doxycycline (many
brand names and
generic)
Prophylaxis in areas
with chloroquine- or
mefloquine-resistant P.
falciparumd
100 mg PO qd (except in
pregnant women; see
Comments)
≥8 years of age: 2 mg/kg PO qd,
up to adult dose
Begin 1–2 days before travel to malarious
areas. Take daily at the same time each day
while in the malarious areas and for 4 weeks
after leaving such areas. Doxycycline is
contraindicated in children aged <8 years and
in pregnant women.
An alternative to
chloroquine for primary
prophylaxis only in the
very few areas with
chloroquine-sensitive
P. falciparumd or areas
with P. vivax only
310 mg of base (400 mg of salt)
PO once weekly
5 mg of base/kg (6.5 mg of salt/
kg) PO once weekly, up to
maximum adult dose of 310 mg
of base
Mefloquine (Lariam
and generic)
Prophylaxis in areas with
chloroquine-resistant
P. falciparumd
228 mg of base (250 mg of salt)
PO once weekly
≤9 kg: 4.6 mg of base/kg (5 mg
of salt/kg) PO once weekly
>9–19 kg: ¼ tablete once weekly
>19–30 kg: ½ tablet once
weekly
>30–45 kg: ¾ tablet once
weekly
>45 kg: 1 tablet once weekly
Begin 1–2 weeks before travel to malarious
areas. Take weekly on the same day of the
week while in the malarious areas and for
4 weeks after leaving such areas. Mefloquine
is contraindicated in persons allergic to this
drug or related compounds (e.g., quinine
and quinidine) and in persons with active
or recent depression, generalized anxiety
disorder, psychosis, schizophrenia, other
major psychiatric disorders, or seizures.
Use with caution in persons with psychiatric
disturbances or a history of depression.
Mefloquine is not recommended for persons
with cardiac conduction abnormalities.
For prevention of malaria
in areas with mainly
P. vivax
30 mg of base (52.6 mg of salt)
PO qd
0.5 mg of base/kg (0.8 mg of
salt/kg) PO qd, up to adult dose;
should be taken with food
Primaquine Terminal prophylaxis to
decrease risk of relapses
of P. vivax and P. ovale
30 mg of base (52.6 mg of
salt) PO qd for 14 days after
departure from the malarious
area
0.5 mg of base/kg (0.8 mg of
salt/kg), up to adult dose, PO qd
for 14 days after departure from
the malarious area
This therapy is indicated for persons who have
had prolonged exposure to P. vivax and/or
P. ovale. It is contraindicated in persons with
G6PD deficiency as well as during pregnancy.